Exoskeleton (pronounced ek-soh-skel-i-tn)
(1) In
zoology, an external covering or integument, any hardened external structure,
as the shells of crustaceans or the scales and plates of fishes and reptiles,
especially when it is of the nature of bone.
(2) All
hard parts, such as hair, teeth, and nails which develop from the ectoderm or
mesoderm in vertebrates (generally used only in technical literature).
(3) A wearable robotic machine used for aided or augmented mobility.
1841: The construct was exo- + skeleton. The prefix endo- was used as a word-forming element meaning "inside, within, internal. It was from the Ancient Greek ἔνδον (éndon) (within; inner; internal) from the primitive Indo-European en-do- (an extended form of the root en (in)). Skeleton was from the New Latin skeleton (bones, bony framework of the body), from the Ancient Greek skeleton soma (dried-up body, mummy, skeleton), from the neuter of σκελετός (skeletós) (dried up, withered, dried body (and as a noun: parched; mummy), from σκέλλω (skéllō & skellein) (dry, dry up, make dry, parch), from the primitive Indo-European (s)kelha- (to parch, wither); related was the Ancient Greek σκληρός (sklirós) (hard). Exoskeleton is a noun; the noun plural is exoskeletons or exoskeleta.
Skelton was an early variant form. The noun use of Greek skeletos passed into Late Latin as (sceletus), hence the French squelette and the rare English skelet (1560s), the Spanish esqueleto & the Italian scheletro. The meaning "bare outline" was first recorded circa 1600; hence the term "skeleton crew" from 1778 used to describe minimal staffing, the skeleton key a similar allusion to some of a structure being removed. The phrase "skeleton in the closet" (source of secret shame to a person, family or institution) is from 1812 and thought an adoption from the imagery in the fable Bluebeard (1697) by Charles Perrault (1628-1703). Exoskeleton was in 1841 coined by by English paleontologist Sir Richard Owen (1804–1892). Exoskeleton has become more widely used in recent years because of the interest in fields such as engineering, robotics and medicine in using external structures, often to augment or replace human functions. As early as the 1960s, the "exoskeleton look" became fashionable among architects who would no longer conceal features like plumbing pipes
Trilobites
Trilobite variants.
Trilobite (pronounced try-low-byte) translates literally as “three lobes". Often casually referred to as bugs or sea-bugs, in taxonomy, all trilobites actually belong in the class of trilobite in the phylum arthropod and within the class are ten orders. It’s not known how many species of trilobites existed but almost 21,000 have thus far been identified in the fossil record, their numbers and variety leading them to be regarded as one of history’s more successful animals. They inhabited all the seas and oceans and endured some three-hundred million years, surviving several mass-extinction events. Their long duration, their structure and living habits meant they became a common and frequently discovered fossil, noted since antiquity although the first attempt scientifically to classify one seems to have been by Wan Shizen of China who, in 1689 described trylobite pygidia (tails) as "batstones". The first known scientific drawing was by Welsh botanist, the Reverend Edward Lhuyd (1660-1709) whose sketch of a trilobite was published in "The Philosophical Transactions of the Royal Society. The drawing, now classified as being a Ogygiocarella debuchii, was then (not unreasonably), called the "flatfish".
Truly ancient, trilobites pre-date the Cambrian explosion (some 540 million years ago) and went extinct only towards the end of the Permian extinction event (some 250 million years ago) which ended the Paleozoic age which had lasted some 290 million years. However, the earlier events took their toll, a few orders vanishing after the Ordovician event (some 445 million years ago) while the Devonian event (some 370 million years ago) removed all but one order, that last survivor dying out in the Permian event. Why such a successful and prolific creature could not endure these extinctions remains a debate, the more popular theories including (1) environmental change happening with such rapidity there was no time for evolutionary adaptation and (2) the needed sources of nutrition vanishing because organisms lower in the food-chain went extinct. All shared the same basic structure, having three lobes: a left pleural, a middle axial and a right pleural lobe, their bodies divided into a cephalon (dead), thorax (middle), and pygydium (tail). Trilobites had a thick, protective exoskeleton which formed a hard calcite shell, something like that of the modern crab and is the reason for their frequency in the fossil record, the exoskeletons usually the only part to survive although, in the rare cases where certain surrounding conditions exist, traces of soft tissue such as antennae can survive fossilization. As a trilobite grew, it molted its exoskeleton, and many of the fossils which exist are molted frames rather than dead creatures.
Before & after avian intrusion: 1952 Mercedes-Benz 300 SL (W194), Carrera Panamericana, Mexico, November 1952.
In nature, exoskeletons evolved over thousands of generations through the interplay of natural selection and adaptation to prevailing environmental conditions but with man-made objects such things, in cases of immediate need, can within hours be fashioned. Returning to the circuits for the first time since the end of World War II (1939-1945), Mercedes-Benz had enjoyed immediate success with the new 300 SL (W194) race car which, despite being underpowered, proved its mettle with a combination of robustness, reliability and outstanding aerodynamics which both increased performance and improved fuel economy. The W194, ten of which were built, is often referred to as “the first gullwing” (although the configuration had in 1939 been seen on a one-off Bugatti and was not uncommon in aviation and nautical use) but the distinctive doors were introduced only after those enforcing the regulations of sports car at Le Mans ruled the 300 SL’s original “doors” were no such things and were merely “elaborately framed windows”. With re-designed doors fitted, the W194 promptly delivered the factory a 1-2 result in the 1952 24 hour endurance classic.
Concluding things that year was the third Carrera Panamericana, a gruelling endurance event of eight stages over some 3,100 km (1,925 miles) between 19-23 November. With typical thoroughness, the factory shipped three W194s along with a large support staff and all was going well until the lead car collided with a large bird (contemporary articles variously reporting the unfortunate creature as a vulture or buzzard) which smashed through the windscreen at an impact speed in excess of 215 km/h (135 mph), stunning the co-driver and leaving him bloodied. Shook awake, he recovered and when the car reached the service point a new windscreen was fitted and, as a precaution against other vultures (or whatever) seeking vengeance, eight metal bars were fitted, the ad-hoc birdcage designed to keep them out, not in. Without further avian “events”, the W194 repeated the result from Le Mans by finishing 1-2 and, fully restored, the winning car is part of the collection of the Mercedes-Benz Museum in Stuttgart.
1968 London-Sydney Marathon: The 1968 Porsche 911S which finished fourth (left) and after a full restoration (right). A zoologist would insist this is only a "semiexoskeleton" on the basis of the partial coverage.
As all in the Northern Hemisphere understand, in Australia, on land, sea & air, all the wildlife will try to kill you. Noting this, when Porsche’s competition department prepared its three entries for the 1968 London-Sydney they limited their concern about sharks and crocodiles to a single paragraph in the drivers’ instruction sheets and decided the main threat to the cars were wandering “giant” kangaroos and low-flying wedge-tailed eagles (a bird with a wingspan wider even than Mexico’s vultures). The first event of its type in the modern era, the 1968 London-Sydney Marathon spanned half the globe over a distance of 12,237 miles (16,934) and the route shows how times have changed, stages in Iran & Pakistan included; neither country now places many organizers of events would include in their itineraries but in 1968, kangaroos were still deemed more a threat than terrorism. Accordingly, the factory designed a kind of exoskeleton, a robust external roll cage which essentially was the W194’s “birdcage” but “on steroids”. As originally conceived, the structure had been of more modest proportions and while the final result may look like “overkill”, things were “beefed up” following one of the drivers writing off a rented VW Beetle after colliding with a kangaroo during pre-event testing in Australia. Impressed by the extent of the damage, the engineers produced an impressively strong protective cage, content that whatever else might go wrong, it wouldn’t be a kangaroo which ended the venture. Porsche 911S #58 finished fourth, victory going to a Hillman Hunter #75, one of the more improbable machines to win an international endurance event.
1968 London-Sydney Marathon: The winning 1968 Hillman Hunter (left) and restored 1967 Pakyan 1725 (right), the Iranian variants often noted for their additional bling. Unlike the cautious Germans, most teams fitted only rudimentary additional protection against kangaroos and such; some ruefully later casting envious glances at Porsche's exoskeleton.
In fairness, though an unlikely competition car, the Hillman Hunter (the “mainstream” model in the “Arrow” range which would sold also as “badge-engineered” Sunbeam, Singer & Humber variants) was a success in many countries, lasting in Europe from 1966 until 1979 although its most extraordinary longevity was achieved in the Islamic Republic of Iran. In 1966, during the reign of Mohammad Reza Pahlavi (1919–1980; last Shah of Iran 1941-1979) assembly from CKD (completely knocked down) packs had begun with the car sold as the Paykan (پیکان, Romanized as Peykān and literally “Arrow”) and unlike many things, it survived the 1979 revolution to continue to flourish under the rule of Grand Ayatollah Ruhollah Khomeini (1900-1989; Supreme Leader, Islamic Republic of Iran, 1979-1989), full local manufacture starting in 1985. Although theologically uncompromising, there’s no record of the Imam having complained of the presence on his assembly lines of a product from شیطان کوچک (Sheytān-e Koochak) (the UK & Israel being “Little Satan” and the US شیطان بزرگ (Sheytān-e Bozorg) “Great Satan” although in the West the terms are often misunderstood because in flavour of Shia Islam practiced in Iran, Satan is a pathetic rather than fearsome figure). When in 1979 Grand Ayatollah Ali Khamenei (1939-2026; Supreme Leader, Islamic Republic of Iran 1989-2026) became supreme leader, the Pakyan carried on and although production of the salon finally ended in 2005, the pick-up variant continued to be sold for another decade, the last not leaving the line until almost 50 years after the first Hunter was sold in the UK. From the modest Hunter, the ayatollah's got their money's worth.
Exoskeleton cars
MVE Exocet (left) & Exomotive’s Exocet Sport V8 (right).
Exoskeleton vehicles are numerous on farms, mine-sites and such but rarely seen on public roads. They do though have a niche for those who want something which sacrifices just about everything (aerodynamics, weather protection, doors etc) for the nimbleness only extreme light-weight can deliver. An example is the MVE Exocet, released for public sale in 2010. It’s an inventive approach to the kit-car concept and takes the classic front-engined, rear-wheel drive approach, based on Mazda’s Miata (the MX-5, introduced in 1989 and a kind of clone of the Lotus Elan of the 1960s but without the problems), the advantage with the Japanese platform being its unusual sub-frame which permits the removal of the body, leaving the engine, drive-train and suspension as a rolling assembly to be transplanted to the Exocet chassis.
Because of the light weight, even when using sensible four-cylinder engines the Exocet delivers high-performance but the Americans in particular can’t resist the idea that just about any car can be improved by the installation of a V8 and quite outlandish power to weight ratios are possible. An indicative example of Exomotive’s Exocet Sport used a 525 horsepower (LS3) version of one of the later evolutions of the small-block Chevrolet V8 which, fully fueled, weighed in at 1690 lb (767 kg, the 2026 Formula 1 regulations set a minimum dry-weight (ie excluding fuel) of 768 kg). Because it possible to buy, off the shelf (as a “crate” engine), V8 engines with about the same power as a F1 power-plant generates, although there was be something a weight penalty, the potential does exist to build a two-seater roadster with a similar power-to-weight ratio and there are jurisdictions which even allow such a thing to be registered for use on public roads. Opinions would differ on whether such a build is a good idea but the little machines, if the V8 was tuned more for low and mid-range torque rather than ultimate power, would seem to have great potential in competitions such as short-course events and hill-climbs although the dubious aerodynamics would render it less suited to high-speed tracks.
Art and money: Porsche 934 (left) by Benedict Radcliffe (b 1976) sold for US$249,002 while Comedian (a banana duct-taped to the wall, right) by Maurizio Cattelan (b 1960) realized US$6.2 million.
The exoskeleton concept inspired English artist and sculptor Benedict Radcliffe to create a number of small scale tubular steel sculptures in the shape of cars including the Lancia Stratos (1973-1978), Lamborghini Countach (1974-1990) and Ferrari F40 (1987-1992), mostly powdered-coated in lurid colors. Usually, they sell for several thousand US dollars but in early 2025, one in 1:1 scale in the shape of a Porsche 934 sold for US$249,002; in a nod to history, it was painted in the same fluro-orange used for the Jägermeister livery used for the race cars in 1979-1977 and rolled on period-correct centre-locking BBS wheels shod with Avon slick tires. At that price, it was little different from what one would pay for a new Porsche 911 GT3, straight from the showroom floor. Still, it’s less than the US$1.5 million which is typical of what’s been paid in recent years on the rare occasions a 934 is offered for sale. Produced between 1976-1977, Porsche built 31 934s, simply for the purpose of creating a version of the 930 (the 911 Turbo, 1975-1978) which would comply with the FIA Group 4 (GT Cars) rules (the 935 was the companion Group 5 (Special Production Cars) project). The art market cannot be assessed with any form of conventional metrics but in paying a quarter-million odd for a tubular structure, one gets quite a lot compared with the Italian visual artist Maurizio Cattelan’s Comedian, (a banana duct-taped to the wall); a couple of months earlier, at auction, it had gone under the hammer for US$6.2 million.
Endoskeleton (pronounced en-doh-skel-i-tn)
(1) In
zoology, the internal skeleton or framework of the body of an animal (generally
the bony or cartilaginous skeleton of vertebrates). Certain invertebrates, such as sponges and
echinoderms, also have endoskeletons.
(2) For
most (non-technical) purposes, a synonym for skeleton.
1838: The construct was endo- + skeleton The prefix exo- was used as a word forming element in words of Greek origin meaning "outer, outside, outer part" and was used from the mid-nineteenth century. It was from the Ancient Greek ἔξω (éxō) (outer; external) and was related to ex (out of). Endoskeleton is used almost exclusively in the biological sciences. For most general purposes, it’s synonymous with skeleton which is the default assumption of use because it’s familiar from humans and most familiar animals. Endoskeleton is a noun; the noun plural is endoskeletons or endoskeleta.
The word endoskeleton may not have been needed had all creatures on earth had “conventional” skeletons like humans, cats, dogs, fish and such; there would have been just “skeletons”. Of course, architects and engineers likely would have been unable to resist coining “exoskeleton” there seems no better word to describe an externally-located superstructure. Both an endoskeleton and exoskeleton are structural frameworks to some degree (sometimes wholly) supporting and shaping an organism’s body and which form an animal evolved to adopt was a product of history and environment. That’s best illustrated by those with hard outer shell (really the ultimate exoskeleton) which functions as a kind of armor-plate. Among man-made objects, both models are used and easily identified although exoskeletons (such as the futtocks in nautical design) probably are more common in anything with a “skin” including buildings, aircraft and ships.
Endoskeleton cars
The Birdcage: The Maserati Tipo 60/61 (chassis #2549).
Endoskeleton cars are far from uncommon but some make the concept more obvious than others. The Maserati Tipo 60/61 (1959-1961) gained the nickname “Birdcage” (by which it’s almost always known) because observers were much taken with the delicacy of the construction. By the late 1950s, space-frames had become familiar to race-car builders but they were usually robust-looking arrangements whereas Maserati had rendered an intricate latticework of some 200 chromoly steel tubes welded often in triangulated form in the points of highest stress, the design delivering both lightness and rigidity.
Mercedes-Benz 300 SLR (W196S, upper) & 300 SL (W198, lower).
One of the reasons the Maserati’s skeleton looked so delicate was that the space-frame had become associated with Teutonic-flavored construction like that used by Mercedes-Benz for its 300 SL & 300 SLR. Both shared the same method of construction but despite the names and the the visual similarity between the two, there were few common components beyond the nuts, bolts & screws. The 300 SL (W198; 1954-1963) was a road car while the SLR (W196S; 1955) was a lengthened version of the W196R Formula One Grand Prix car with a sexy body and an enlarged (though somewhat detuned) straight-eight engine. Despite appearing much more substantial than the Maserati's birdcage, the German space-frame was remarkably light.










